At the highest level, Rails requests are served through an application server, which is responsible for directing an incoming request into a Ruby process. Popular application servers that use the Rack web request interface include Phusion Passenger, Mongrel, Thin, and Unicorn.
Rack parses all request parameters (as well as posted data, CGI parameters, and other potentially useful bits of information) and transforms them into a big Hash (Ruby’s record / dictionary type). This is sometimes called the env hash, as it contains data about the environment of the web request.
In addition to this request parsing, Rack is configurable, allowing for certain requests to be directed to specific Rack apps. If you want, for example, to redirect requests for anything in your admin section to another Rails app, you can do so at the Rack level. You can also declare middleware here, in addition to being able to declare it in Rails.
Those requests that are not directed elsewhere (by you in Rack) are directed to your Rails app where it begins interacting with the Rails ActionDispatcher, which examines the route. Rails apps can be spit into separate Rails Engines, and the router sends the request off to the right engine. (You can also redirect requests to other Rack compatible web frameworks here.)
Once in your app, Rails middleware – or your custom middleware – is executed. The router determines what Rails controller / action method should be called to process the request, instantiates the proper controller object, executes all the filters involved, and finally calls the appropriate the action method.
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